Annulus (Geissleria)

An annulus (in genus Geissleria) is a group of apical striae separated from the regular striae of the valve face. An annulus has one to four transapical striae that interrupt the typical striae on each side of the distal raphe ends. The annulus may be separated from typical striae by a narrow hyaline area which may have a ring shape. From Latin for ring. Plural is annuli.

The areolae in these striae may be larger, more elongate, and more densely spaced than the rest of the areolae. Internally in some species, these areolae possess a cluster of silica outgrowths.

Annuli may be distinct or barely discernible, depending on the species. For example, the annuli of G. acceptata are difficult to resolve in LM.

This group of striae was named for the hyaline ring around them in Navicula annulata (now Geissleria annulata). This feature separates Geissleria from Navicula.

annulus-geissleria-lm
Image Credit: E.F. Stoermer
A Geissleria species in valve view (LM). Arrow indicates one of the stria in this annulus.
annulus-geissleria-cascadensis
Image Credit: Rosalina Stancheva
Internal view of Geissleria cascadensis (SEM). The annulus shown here consists of three striae on either side of the distal end of the raphe. The areolae of this annulus are covered by silica outgrowths. Scale bar = 2 µm.
annulus-geissleria-sem1
Image Credit: M. Potapova
Internal view of Geissleria acceptata (SEM). Arrow indicates an areola in this annulus. This areola has silica outgrowths around the opening.