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The fistula is an isolated tube, or pore near the valve central area. The fistula has a single slit-like external opening. The structure opens to the valve interior by either a single circular opening, or a transapical linear array of up to five circular to elliptic openings. Internally, the fistula is domed and hymenate.
A fistula is present in the genera Proschkinia and Fistulifera. The structure is analogous to the stigma (but not homologous).