The ratio of mantle height to valve diameter is either equal to, or greater than, 1. On the valve mantle, two diametrically opposed grooves containing no areolae extend the entire length of the valve and two large, pointed separation spines diametrically opposed on the valve face extend in the plane of the mantle onto grooves of the neighboring valves. Mantle reolae may not be visible in LM. Valves lack linking spines. The ringleiste is solid and relatively shallow.