Steve was raised near the sand dune successional communities of Lake Michigan in northwestern Indiana. He has a B.S. and M.A. from Valparaiso University and then taught secondary biology and chemistry in Mill City, Oregon. Steve felt drawn to marine biology, though, and furthered his education by obtaining a doctorate from Oregon State University studying estuarine epiphytic diatoms with C. David McIntire. He is now retired after 34 years of undergraduate biology teaching at Wartburg College, Waverly, Iowa. Steve continues to study the diatoms, including species of the Cedar River (part of the Iowa-Cedar basin), fens of Iowa and Minnesota, freshwater and intertidal zones of Hawaii, and recovering wetlands of central Iowa.

Species contributed

Craticula ambigua | Diprora haenaensis

Genera contributed